I think Guideline 1 is more or less for those who really want to try and simulate a Star Trek universe experience. I regret that many people would feel locked into a role that limit their creativity and game play style.
The point to GL1 is that people should keep RP in mind when playing the race. IE the Borg do not normally forge alliances with the Federation and then conquer the galaxy. But that being said, it really is up to the player.
I would hope that a player would appreciate the limitations of his/her race and play them accordingly, but that is really an unobtainable expectation given the limiting nature that such arbitrary boundaries would generate. A player must be free to play his or her race the way they deem necessary for their game goals.
I agree that Role Playing is important, but everyone that has ever played a game knows that treaties are necessary for survival. The problem here is that some races are defined as out right hostile. The Borg and 8472 being the examples. However, 8472 and the Borg have both allied themselves with the Federation on at least one occation each.
So treaties between HOSTILE natured players is a natural evolution of game play regardless of race characteristics.
Besides; "Make the enemy of thy enemy thy friend, then on the eve of victory against our common foe, attack thy ally and assure utter and complete victory for yourself and yourself alone."